Haplogroup E3b1a2 as a Possible Indicator of Settlement in Roman Britain
by Soldiers of Balkan Origin
Abstract
The invasion of Britain by the Roman
military in CE 43, and the subsequent occupation of Britain for nearly four
centuries, brought thousands of soldiers from the Balkan peninsula to Britain
as part of auxiliary units and as regular legionnaires. The presence of Haplogroup E3b1a-M78 among
the male populations of present-day Wales, England and Scotland, and its nearly
complete absence among the modern male population of Ireland, provide a
potential genetic indicator of settlement during the 1st through 4th Centuries
CE by Roman soldiers from the Balkan peninsula and their male Romano-British
descendants. Haplotype data from several
major genetic surveys of Britain and Ireland are examined, analyzed and
correlated with historical, epigraphic and archaeological information, with the
goal of identifying any significant phylogeographic associations between
E3b1a-M78 and those known Romano-British settlements and military posts that
were associated specifically with Roman soldiers of Balkan origin. Studies by Cruciani et al. (2007), Perečić et
al. (2005), and Marjanovic et al. (2005), examining the distribution of
E3b1a-M78 and E3b1a2-V13 in the Balkans, are analyzed further to provide
evidence of phylogeographic associations between the E3b1a2 haplotypes
identified within the Balkans by these studies and those regions of the Balkans
occupied first by the Roman army in antiquity.
E3b1a2 is found to be at its highest frequency worldwide in the
geographic region corresponding closely to the ancient Roman province of Moesia Superior, a region that today
encompasses Kosovo, southern Serbia, northern Macedonia and extreme
northwestern Bulgaria. The Balkan
studies also provide evidence to support the use of E3b1a-M78 (in the present
study) as a close proxy for the presence of E3b1a2-V13 (representing 85% of the
parent E3b1a-M78 clade) in both the Balkans and in
Address for
correspondence: stevenbird1000@hotmail.com
Received:
Introduction
The
origins, arrival times and possible routes of migration of the E3b haplogroup[1]
to
At about
the same time as the release of Oppenheimer’s book, Bryan Sykes (2006)
published his book Blood of the Isles
(entitled Saxons, Vikings, and Celts
in the
Very
shortly after the publication of these two books, Cruciani et al. (2007)
published a new study defining ten subclades of haplogroup E3b1a-M78 through
several newly identified unique event polymorphisms (UEP’s).[7]
The subclade E3b1a2 (identified by the presence of the V13 and V36 UEPs) was
found by Cruciani et al. (2007) to have a strong phylogeographic association
with the southern Balkan peninsula; this subclade also was found by the same
study to correspond very closely to the α (“alpha”) cluster of E3b1a-M78,
first identified by Cruciani et al., (2004) using microsatellite (
Semino et
al. (2004) viewed E3b1a-M78, of which E3b1a2 is, by far, the most common
subclade in Europe, as an indicator of the diffusion of people from the Balkans
(along with a “companion” clade, J2b1-M12/M102) and therefore a candidate for a
residual genetic signature of the Neolithic demic diffusion model. Cruciani et
al. (2007) have brought the Neolithic dating assumption into question, however,
by their revised dating of the expansion of E-V13 and J-M12, from the Balkans
to the remainder of
Two
dating methods were employed by Cruciani (2007) to calculate the “time to most
recent common ancestor” ("TMRCA"): that of Zhivotovsky et al. (2006)
based on his “evolutionary effective” mutation rate for an average square
distance ("
An
important finding of this study was that E-V13 and J-M12 had essentially
identical population coalescence times.
They concluded that the E-V13 and J-M12 subclades expanded in
Our estimated coalescence age of about 4.5
ky for haplogroups E-V13 and J-M12 in Europe (and their C.I.s) would also
exclude a demographic expansion associated with the introduction of agriculture
from Anatolia and would place this event at the beginning of the Balkan Bronze
Age, a period that saw strong demographic changes as clearly testified from
archeological records.
These
expansion times were calculated by Cruciani (2007) to have occurred between
4.0-4.7 kya for E-V13 and 4.1-4.7 kya for J2-M12, with the upper limit of the
expansion time for E-V13 at 5.3 kya and for J2-M12 at 6.4 kya. Both expansion times therefore are centered
at approximately 4.3-4.35 kya, a period of time corresponding to the EBA in the
southern Balkans (Hoddinott, 1981).
Cruciani
et al.’s E-V13 and J2-M12 coalescence times bear a striking similarity to
carbon-14-based date calculations for certain archaeological sites in the
Maritsa river valley and its tributaries, near the city of Nova Zagora,
Bulgaria (Nilolova, 2002). These sites
are associated directly with the proto-Thracian culture of the southern Balkans
that came to dominate the region during the first millennium
Another
difficulty for the acceptance of Oppenheimer’s “Neolithic” arrival time for E3b
and J2 in
If
E3b1a-M78 had in fact arrived during the Neolithic era by water routes from
While a
Neolithic arrival date for E3b in
Bronze Age and Iron Age Celtic-style cultures have been
identified by both Oppenheimer and Sykes as being associated with the so-called
"Western Atlantic Modal Haplotype" (R1b1c). Alcock (1972, pp. 99-112) has examined the
model of a Celtic Irish-Sea culture-province in the pre-Roman Iron Age (“IA”),
in particular connections across the Irish Sea, including a dominant Irish
cultural component, as well as related settlement in Wales, Strathclyde,
Argyll, and southwestern Britain. The
problems encountered by the Neolithic theory of Oppenheimer, i.e., the virtual
absence of E3b and the complete absence of J in
Difficulties
with Neolithic, BA, and IA models for the migration of E3b and J2 to
Methodology
The three
data sets of Capelli, Weale, and Sykes used six to ten
A factor
preventing direct comparison (by percentage) of these three data sets was a
substantial difference in the format used for reporting the geographic origins
of individual haplotypes. Weale (2002)
and Capelli (2003) each specified geographic locations (towns or villages) in
their reports of haplotype frequencies; Sykes (2006), however, reported his
findings by combining locations into larger geographic regions that, in most
cases, joined several British counties into a single data set (such as "
Even with
these limitations, however, it was possible to identify some trends in the
three data sets when combined. Using
|
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|
|
|
|
|
|
|
|
|
Figure 1. Histograms of E3b-M35, Allele Frequencies by
Subclade
As a
third method of estimation (and to provide a check of the robustness of the
derived subclade assignments), a median-joining network was constructed in NETWORK
4.2.0.1,[14]
using the allele data from the OGAP predicted as E3b by Athey's Y-Haplogroup
Predictor, as shown in Figure 2.[15] The E3b (estimated) data from the OGAP is
presented in Table 1, along with the subclade assignments determined
using the methodology outlined above.
These assignments are also presented in Table 2, grouped
according to the OGAP's geographic regions and the overall percentages for each
E3b subclade.

Key to multiple taxa nodes:
Modal: A2960, 5371, 4018, A3040, A3097, A3174,
A2923, A2833
A3429, A2967,A3029,
A9065 (E-M78)
2745: 2745,
A2243, A2950, A1201 (E-M78)
738: 738,
A2090, A2109, A3093 (E-M78)
5251: 5251, A8115, A8135, A8584
(E-M78)
A2981: A2981, 503 (E-M78)
A2751: A2751, A2211 (E-M78)
A2547: A2547, A231 (E-M78)
(All
other nodes have one taxon each.
Geographic descriptions apply only to the adjacent taxon).
Figure 2. Median-Joining Network, OGAP Data, E3b
Table
1. OGAP E3b Data Grouped by Region and
Classified by Subclade
|
|
|
3 |
3 |
1 |
3 |
4 |
3 |
3 |
3 |
3 |
4 |
|
|
||
|
|
|
9 |
9 |
9 |
9 |
2 |
8 |
8 |
9 |
8 |
2 |
|
|
||
|
OGAP
|
|
3 |
0 |
|
1 |
6 |
8 |
9 |
2 |
9 |
5 |
|
|
||
|
Haplotype
|
|
|
|
|
|
|
|
| |
|
| |
|
Estimated |
|
||
|
Number |
OGAP
Regional Identification |
|
|
|
|
|
|
1 |
|
2 |
|
Subclade |
|
||
|
738 |
Argyll
|
13 |
23 |
13 |
10 |
|
|
13 |
11 |
17 |
0 |
E3b1a-M78 |
|
||
|
A2547
|
Borders |
13 |
24 |
13 |
10 |
11 |
12 |
14 |
11 |
17 |
0 |
E3b1a-M78 |
|
||
|
A2960
|
|
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
12 |
E3b1a-M78 |
|
||
|
5251 |
|
13 |
24 |
14 |
10 |
|
|
13 |
11 |
18 |
|
E3b1a-M78 |
|
||
|
5371 |
|
13 |
24 |
13 |
10 |
|
|
13 |
11 |
17 |
|
E3b1a-M78 |
|
||
|
5924 |
|
13 |
24 |
13 |
9 |
|
|
14 |
11 |
16 |
|
E3b1b-M81 |
|
||
|
3512 |
Grampian
|
13 |
24 |
13 |
10 |
|
|
13 |
11 |
16 |
|
E3b1a-M78 |
|
||
|
4018 |
Grampian
|
13 |
24 |
13 |
10 |
|
|
13 |
11 |
17 |
|
E3b1a-M78 |
|
||
|
4022 |
Grampian
|
13 |
24 |
13 |
10 |
|
|
13 |
11 |
19 |
|
E3b1a-M78 |
|
||
|
2745 |
|
13 |
24 |
13 |
10 |
|
|
13 |
11 |
18 |
|
E3b1a-M78 |
|
||
|
A2243
|
|
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
18 |
0 |
E3b1a-M78 |
|
||
|
A3040
|
|
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
10 |
E3b1a-M78 |
|
||
|
A3097
|
|
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
12 |
E3b1a-M78 |
|
||
|
A3359
|
|
13 |
24 |
13 |
10 |
11 |
12 |
12 |
11 |
17 |
0 |
E3b1a-M78 |
|
||
|
A2689
|
|
15 |
25 |
13 |
11 |
11 |
12 |
11 |
11 |
19 |
12 |
E3b1c1-M34 |
|
||
|
A2090
|
|
13 |
23 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
12 |
E3b1a-M78 |
|
||
|
A2109
|
|
13 |
23 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
0 |
E3b1a-M78 |
|
||
|
A3093
|
|
13 |
23 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
12 |
E3b1a-M78 |
|
||
|
A2112
|
|
13 |
24 |
15 |
10 |
11 |
12 |
13 |
11 |
18 |
12 |
E3b1a-M78 |
|
||
|
A3174
|
|
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
12 |
E3b1a-M78 |
|
||
|
A705 |
|
13 |
24 |
13 |
11 |
11 |
12 |
12 |
11 |
17 |
12 |
E3b1a-M78 |
|
||
|
A2923
|
|
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
0 |
E3b1a-M78 |
|
||
|
A8115
|
Shetland |
13 |
24 |
14 |
10 |
|
|
13 |
11 |
18 |
|
E3b1a-M78 |
|
||
|
A8135
|
Shetland |
13 |
24 |
14 |
10 |
|
|
13 |
11 |
18 |
0 |
E3b1a-M78 |
|
|
|
|
A8584
|
Shetland |
13 |
24 |
14 |
10 |
|
|
13 |
11 |
18 |
0 |
E3b1a-M78 |
|
||
|
A2981
|
|
13 |
23 |
13 |
10 |
11 |
12 |
13 |
11 |
18 |
0 |
E3b1a-M78 |
|
||
|
A2629
|
|
14 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
0 |
E3b1a-M78 |
|
||
|
A2751
|
|
13 |
24 |
14 |
10 |
11 |
12 |
13 |
11 |
17 |
0 |
E3b1a-M78 |
|
||
|
A2833
|
|
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
10 |
E3b1a-M78 |
|
||
|
A2950
|
|
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
18 |
10 |
E3b1a-M78 |
|
||
|
A3429
|
|
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
13 |
E3b1a-M78 |
|
||
|
A3059
|
|
13 |
24 |
13 |
10 |
11 |
12 |
11 |
11 |
17 |
10 |
E3b1a-M78 |
|
||
|
A2581
|
South-west |
13 |
23 |
13 |
10 |
11 |
12 |
15 |
13 |
16 |
12 |
E3b1-M35 |
|
||
|
A2663
|
South-west |
14 |
24 |
13 |
10 |
11 |
12 |
13 |
13 |
16 |
12 |
E3b1-M35 |
|
||
|
A2967
|
South-west |
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
13 |
E3b1a-M78 |
|
||
|
A3029
|
South-west |
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
17 |
10 |
E3b1a-M78 |
|
||
|
A3153
|
South-west |
13 |
25 |
14 |
10 |
11 |
12 |
13 |
11 |
18 |
0 |
E3b1a-M78 |
|
||
|
A3165
|
South-west |
13 |
24 |
13 |
10 |
11 |
12 |
14 |
11 |
19 |
12 |
E3b1a-M78 |
|
||
|
503 |
Strathclyde
|
13 |
23 |
13 |
10 |
|
|
13 |
11 |
18 |
|
E3b1a-M78 |
|
||
|
359 |
Strathclyde
|
13 |
25 |
13 |
10 |
|
|
13 |
11 |
17 |
|
E3b1a-M78 |
|
||
|
A9065
|
Tayside & Fife |
13 |
24 |
13 |
10 |
|
|
13 |
11 |
17 |
|
E3b1a-M78 |
|
||
|
A1201
|
|
13 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
18 |
0 |
E3b1a-M78 |
|
||
|
A231 |
|
13 |
24 |
13 |
10 |
11 |
12 |
14 |
11 |
17 |
12 |
E3b1a-M78 |
|
||
|
A2211
|
|
13 |
24 |
14 |
10 |
11 |
12 |
13 |
11 |
17 |
0 |
E3b1a-M78 |
|
||
|
A1335
|
|
14 |
24 |
13 |
10 |
11 |
12 |
13 |
11 |
15 |
0 |
E3b1-M35 |
|
||
Table 2. E3b Subclades by OGAP Region and Percentage
|
Subclade |
Region |
N=(total) |
n= |
% |
|
|
|
|
M-78 |
Argyll
|
19 |
1 |
5.3% |
|
|
|
M-78 |
|
163 |
7 |
4.3% |
|
|
|
M-78 |
|
151 |
4 |
2.6% |
|
|
|
M-78 |
|
125 |
3 |
2.4% |
|
|
|
M-78 |
Grampian
|
126 |
3 |
2.4% |
|
|
|
M-78 |
Tayside & Fife |
56 |
1 |
1.8% |
|
|
|
M-78 |
South-west |
237 |
4 |
1.7% |
|
|
|
M-78 |
|
178 |
3 |
1.7% |
|
|
|
M-78 |
Strathclyde
|
120 |
2 |
1.7% |
|
|
|
M-78 |
|
365 |
6 |
1.6% |
|
|
|
M-78 |
Borders |
64 |
1 |
1.6% |
|
|
|
M-78 |
|
82 |
1 |
1.2% |
|
|
|
M-78 |
Northern Isles |
202 |
1 |
0.5% |
|
|
|
|
Total |
1888 |
37 |
2.0% |
|
|
|
|
|
|
|
|
|
|
|
M-81 |
|
125 |
1 |
0.8% |
|
|
|
|
Total |
125 |
1 |
0.8% |
|
|
|
|
|
|
|
|
|
|
|
M-34 |
|
151 |
1 |
0.7% |
|
|
|
|
Total |
151 |
1 |
0.7% |
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
M-35 |
South-west |
237 |
2 |
0.8% |
|
|
|
M-35 |
|
178 |
1 |
0.6% |
|
|
|
|
Total |
415 |
3 |
0.7% |
|
|
|
|
|
|
|
|
|
|
|
n/a |
|
193 |
0 |
0.0% |
|
|
|
|
Total |
193 |
0 |
0.0% |
|
|
|
|
|
|
|
|
|
The
locations of individual E3b haplotypes identified on the main
Forty
haplotypes from the OGAP were classified as E3b1-M78 (88.8%). Of the other three subclades predicted, two
E3b*-M35 haplotypes were found in "
Haplogroups
were named according to ISOGG (2007).
E3b Patterns of Settlement in
The frequency
of E3b in Britain was observed to be most prevalent in two regions; a
geographic cluster of haplotypes extending from Wales eastward to the vicinity
of Nottingham, encompassing the region surrounding Chester, and a second (NNE
to SSW) cluster extending from Fakenham, Norfolk to Midhurst, Sussex. Other E3b data points were scattered widely
throughout
|
Figure 3. Frequency of E3b Grouped by County |
Figure 4. E3b Distribution by Geographic Location |
|
Figure 5. E3b Distribution by Frequency and Location |
Figure 6. E3b Distribution– Kriging Method |
Of
special interest is a void appearing in the geographic distribution of E3b,
found largely in the part of
The major
E3b1a-M78 (
The Central English-North Welsh barrier cannot be
explained purely as a simple isolation-by-distance phenomenon because it
contrasts strongly with the lack of evidence for a cline among the five widely
separated English towns. Our findings
are particularly striking, given the high resolution and rapid mutation rate of
the Y chromosome haplotypes on which they are based. These allow genetic barriers, if they exist,
to be clearly defined.
The best explanation for our findings is that the
Anglo-Saxon cultural transition in
The data reported by the OGAP concerning the
"Eshu" void, when combined with Capelli and Weale, provides some
additional evidence that appears to support Weale's hypothesis. The distinct clustering of E3b1a-M78 from
An alternative interpretation of
the Weale and Capelli findings, hypothesized by Thomas et al. (2006), was that
the genetic admixture of
Balkan
Ethnogenesis
The
hypothesis that members of the E3b1a2 haplogroup migrated to
In Saxons, Vikings and Celts, Sykes (2006,
p. 224) queried:
But who were the soldiers of the Roman army? Not all from
Thracian
and Dacian soldiers originating from the geographic regions near the
With the
six main subclades of E-M78 separated into their component parts, according to
Cruciani (2007) it appeared that the frequency distribution of E-V13 was
centered in the region of

Figure 7. E-V13 Frequency, Balkan Region, after
Cruciani et al. 2007, Fig. 2D.
The
E-M78α cluster of E3b1 was found to be identified closely with the
southern Balkans, in particular the regions of the
Figure
8 shows the
distribution of E-V13 in the Balkans with the two data sets combined and
averaged, and with Kosovo treated as a distinct region from
|
Figure 7. E-V13 Frequency, Balkan Region, after
Cruciani et al. 2007, Fig. 2D. |
Figure 8. E-V13 Frequency Contour (Kriging Method),
Southern Balkans. |
|
Figure 9. E-M78 Frequency Contour (Kriging Method),
Southern Balkans. |
Figure 10. Boundary of |
Moesia Superior was roughly rectangular in shape,
with the
. . . a general conclusion may be permitted, that the
original inhabitants of
The
Dacian kingdom, immediately north of the Danube and Moesia Superior, was conquered
partially by Trajan in CE 106, with the region conquered becoming the Roman
province of Dacia Traiana. "Dacian" was a Roman ethnic label;
previously known to the Greeks as the Getae,
Wilcox (1982), Hoddinott (1981), and Webber (2001) have all identified the
Geto-Dacians as people of proto-Thracian descent and relationship. Because of later difficulties with the Goths,
The
Russian linguist Georgiev (1960) stated that the modern Albanian people were
descended from Daco-Mysian ancestors, who had occupied ancient homelands in
western
Thracian soldiers in Roman
Britain
Epigraphic
evidence for the presence of individual Thracian soldiers, as well as for
Thracian military units of the Roman army, is found in several locations in
Sources
for these attestations include bronze military diplomata (documents issued to retiring soldiers) and epigraphic
inscriptions, typically tombstones or dedications. The diplomata, which functioned as retirement papers, also granted
Roman citizenship to retiring soldiers after twenty-five years of service. An
example, found in 1812 near Malpas,
The
inscription is translated (Collingwood, 1990) as follows:
The Emperor Caesar Nerva Traianus Augustus, conqueror of
Germany, conqueror of Dacia, son of the deified Nerva, pontifex maximus, in his
seventh year of tribunician power, four times acclaimed Imperator, five times
consul, father of his country, has granted to the cavalrymen and infantrymen
who are serving in four alae and eleven cohorts called: (1) {ala} I Thracum and (2) I
Pannoniorum Tampiana and (3) Gallorum Sebosiana and (4) Hispanorum Vettonum,
Roman citizens; and (I) {cohor} I Hispanorum and (2) I Vangionum, a thousand
strong, and (3) I Alpinorum and (4) I Morinorum and (5) I Cugernorum and (6) I
Baetasiorum and (7) I Tungrorum, a thousand strong, and (8) II Thracum and
(9)
Among the
units listed in this document, several can be identified with regions found to
be high in frequency for E3b1a2. The ala I Thracum (First Wing of Thracian
Cavalry) and the cohors II Thracum
(Second Thracian Infantry) would have had members from the regions containing
Thracian tribes, including the Roman provinces of Thracia and Moesia. The
Pannonian and Dalmatian units, ala I
Pannoniorum Tampiana, and the cohors
IIII Delmatarum (Fourth Dalmatian Infantry) may have had some E3b1a2
members from the Balkan peninsula also, although clearly Reburrus himself (for
whom this particular diploma was issued) or his father was “from Spain.”
Frere et
al., have stated that “less than 20% of diploma recipients moved out of the
province in which they had served upon retirement.”[27]
It is evident from the
wording of the Malpas diploma that many of the soldiers must have had wives and
children, who stood to gain rights as citizens upon the soldiers’
retirement. The fact that this bronze
plaque was found barely twenty miles (32 km) from the major fortress at Deva (now Chester, Cheshire), not far
from the Roman Road and very close to the minor Roman settlement of Bovium
(now Tilston, Cheshire) would tend to suggest that Reburrus settled in the
immediate vicinity following his retirement.
Presumably, soldiers from retiring Thracian, Dalmatian or Pannonian
units, all of which were likely to have had members whose haplotype was E3b1a2,
would have done the same.
Another
diploma, which was discovered in Middlewich,
Aside
from the Malpas and Middlewich diplomas, a considerable amount of epigraphic
evidence has been identified linking Roman soldiers who were buried in Deva
specifically to the Roman provinces of Thracia and Moesia, including in some
cases place names of origin within these provinces for these soldiers
(Carrington, 2006). Four individuals
from the Legio II Adiutrix were
identified by Carrington (2006) from burial markers as having originated in Aprus (now Apri, in Turkish Thrace) and
were associated with the Roman fortress at Deva in the period from CE 71 to
circa 86. There is also evidence of
soldiers of Thracian origin who were members of the XX Legio VV (“Twentieth Legion Valeria Victrix”) settling in or
near the Roman fortress of Deva (now Chester, Cheshire), presumably upon
retirement (Carrington, 2006). A third
century grave marker depicts a soldier from the XX Legio VV and his spouse, identifying him as being “Bessus”
(of the tribe of the Bessi).[29]
This stone is depicted in The Roman
Inscriptions of Britain ("RIB") entry
no. 523 (Collingwood, 1965, p. 174). The
find is significant because it provides direct epigraphic proof of a married
Thracian soldier who lived (and died) in Deva during the third century.[30] The description from the RIB is as follows:
523. Tombstone, 25
X 46 in., broken across the stool and also above the inscription. Donatus reclines on a couch, and holds in his
left hand a scroll, probably his will, and in his right a cup. On the left his wife reclines behind
him. Found in 1887 in the North Wall
(east part). Now in the
[Inscription]: D(is) M(anibus) | C(a)ecilius Donatus B |
essus na | tione mili | tauit ann | os XXVI uix | it
annos XXXX.
To the spirits of the departed, Caecilius Donatus, a
Bessian tribesman, served 26 years and lived 40 years.
The Bessi were a Thracian tribe. Since Donatus given no
praenomen, tribe, or father’s names and uses the spelling Cecilius (for
Caecilius), and as a legionary was recruited from a tribal area and not from a
town, a third-century date seems certain.
Mócsy
(viz., quotation above) identified a third century tombstone in which a soldier
from Scupi (
Carrington
(2006, p. 12) also addressed the evidence for the Chester canabae (civilian settlements), including marriages between Roman
soldiers and local women, the legal positions of the soldier, his wife and
their offspring with regard to Roman citizenship and inheritance, and the most
probable period of settlement:
The first half of the third century is the period when one
would expect family-formation at
Archaeological
evidence for Roman settlement in northern
They are associated, also, with terraced fields which have
resisted destruction even more strongly than the homesteads themselves. Further, sufficient sites have been excavated
to justify the presumption that the majority, if not all, are of the Roman
period.
Hogg
(1965, p. 33) hypothesized that these homesteads associated with the terraced
fields were, in all likelihood, of early third century Roman origin and stated
that:
. . . some proportion of the enclosed homesteads represent
officially encouraged new settlement superimposed on relatively sparse
occupation which had persisted from before the Roman conquest, possibly
enfeebled by punitive measures.
He
estimated the Romano-British population of the region at 4000, plus another 500
soldiers stationed at Segontium (Caernarfon,
Gwynedd). In northeast Wales, the
settlement of Dinorben (1.8 miles
[2.9 km] southeast of Abergele, Clwyd), which was established by the Romans at
the site of an earlier, aba
Abergele
was discovered by Weale (2002) to have the highest percentage of E3b in
In
addition to the tight clustering of E3b1a-M78 haplotypes in Wales and Northern
England (Chester to Nottingham), another significant cluster of E3b1a-M78
appears to follow the coastline of the North Sea and the English Channel from
Fakenham, Norfolk to Midhurst, Sussex (see Figure 6). It could be argued reasonably that the
substantial presence of E3b1a-M78 in a region contained entirely within the
early medieval
More
significant may be the close geographic proximity of some of these E3b1a-M78
haplotypes to the Roman fortresses and settlements known collectively as the
“Forts of the
In Rome’s Saxon Shore Fields (2006, p. 38)
stated that under Severus Alexander (r. 222-235) it was thought advisable to
withdraw troops from the northern frontier (i.e., Hadrian’s Wall) and place
them in new forts built at Branoduno
(Brancaster), Caister-on-Sea, and Regulbium
(Reculver). Epigraphic evidence of
the
Through
aerial photography, it has been determined that most of these forts had small,
non-military settlements (vici) adjacent
to them, with the most extensive being at Branodunum
(Brancaster) (Fields, 2006, p. 51).
From the third century onward soldiers were permitted to have their
families stationed with them (Fields, 2006, p. 48). Moreover, the presence of Roman soldiers,
originating from the Balkans, in the forts of the
Branodunum
was the permanent base of the Equites
Dalmatae Branodunenses (Dalmatian Horsemen of Branodunum). Likewise, the cuneus equitum Dal-matarum Fortensium
(Formation of Brave Dalmatian Horsemen) were stationed at Bradwell, Essex (Othona) (Fields, 2006). The Roman

Figure 11. "
E3b Haplotype Locations (from
Figure 4)
and E3b (Kriging Method) Gradient.
There also
is substantial archaeological evidence for dense settlement by the
Romano-British in the region known today as
Longinus Sdapeze |
matygi f(ilius) duplicarius |
ala prima
Tracum pago |
Sardi(ca)
anno(rum) XL aeror(um) XV |
heredes exs
testam(ento) [f(aciendum)] c(urauerunt |
h(ic) s(itus) e(st)
The translation
in the RIB has some difficulties with the last name of Longinus, identifying it
(probably in error) as “Sdapezematygus,” but also acknowledging that Die Thrakische Sprachreste (
Following this,
the translation becomes:
Longinus
Sdapeze, son of Matygus, duplicaris from the First Cavalry Regiment of
Thracians, from the district of Sardica [Serdica], age 40, of 15 years’
service. His heirs under his will had
this set up. He lies here.
Notwithstanding
the apparent difficulties of the father’s and son’s names, there is a
considerable amount of information to be gleaned from this stone. Longinus was from the region (“pago”)
surrounding Serdica (now
The figure
depicted on the stone is interesting, with obvious similarities to the Thracian
“Hero” religious icon described at length by R. F. Hoddinott (1981, pp.
179-185) in his landmark archaeological treatise, The Thracians. The RIB described the stone figure as follows:
The cavalryman . . . is in
scale-armour, with oval shield on left arm, and is riding to the right on a
richly caparisoned horse, beneath whose belly crouches a naked barbarian with
his left hand holding his shield under him.
In the right hand of the cavalryman there is a dowel-hole, presumably
once filled by a metal weapon [N.B., probably a lance; see the description for
the stone of Rufus Sita, below].
The
mounted Thracian “Hero” figure was a common theme for tombstones from Thracian
units, though not always associated exclusively with ethnic Thracian
soldiers.
Dark
(2002, pp. 97-100) has theorized that sub-Roman Essex may have survived intact
until the sixth century and that the civilian authority may have transitioned
smoothly from sub-Roman to Saxon authority without any evidence of struggle or
the displacement of the local Romano-British population. Drury and Rodwell (1980, p. 71) have provided
similar evidence for the survival of sub-Roman Essex well into the Anglo-Saxon
period. At Rivenhall, two large domestic
buildings and an aisled barn were modified during the 4th, 5th and 6th
centuries, "to suit the needs of the agriculturally-based estate . . .
there is good reason at least to ask whether Roman estates did not gradually
transmute into Domesday vills."
The
authors also stated (Drury, 1980, pp. 71-74):
With the exception of a small number of early Saxon
settlements founded in the coastal areas of Essex, of which Mucking is the best
known, it has become clear that there cannot have been any great influxes of
Germanic immigrants in the 5th and 6th centuries,
comparable to those of East Anglia or east Kent. This is not to say that folk of Germanic
origin were not present in the hinterland of
This
position was echoed by Bassett (1989, p. 25), who stated:
Elsewhere there may have been a takeover (by new
immigrants or by a settled community nearby) of a former Romano-British town
and its immediate hinterland... A good example is provided by the small walled
Roman town at Great Chesterford on the north-western border of the
Peterson
(2004, Fig. 3, p. 66) has described evidence of several Roman cadastres or "centuriations"
in southeastern
The area
north of Saffron Walden and east of the Roman fort at Great Chesterford included
several villas and the largest group of Roman burial tumuli in western Europe,
located at Bartlow, just over the Cambridgeshire border from
An
illustration from 1845 by Knight (1845, Fig. 21, No. 18) provided an interior
view of the largest of the tumuli, a sketch of the burial gallery of tumulus
no. 3, accessed from the exterior originally by a hallway-like passage. In The
Thracians, Hoddinott (1981, pp. 119-121, 124-126) described earthen burial
tumuli very similar in appearance to the Bartlow tumuli, complete with hallways
and burial galleries, throughout ancient Thracia. The Bartlow tumuli are also
similar in appearance to a line of Roman barrows found in
The
tumuli (originally in
Charles
Thomas (1966, pp. 74-98) has commented on the difficulty of archaeological
research in Dumnonia (Cornwall and
Devon) and that due to the material poverty of the region, both in the
pre-Roman and Roman eras, few artifacts have been recovered. He identified the Roman fort of Durocornavis at the (modern) location
called "The Rump," fifteen miles (24 km) southwest from Tintagel,
near Polzeath,
The tombstone of
Rufus Sita, very similar to that of Longinus Sdapeze, is described in RIB
121. It was found in 1824 in Wotton,
about 0.75 miles (1.2 km)
Rufus
Sita eques c(o)ho(rtis) VI
Tracum
ann(orum) XL stip(endiorum) XXII
heredes exs test(amento)
f(aciendum) curaue(runt)
h(ic) s(itus) e(st)
Translated:
Rufus Sita, trooper of the Sixth
Cohort of Thracians, aged 40, of 22 years’ service. His heirs had this erected according to the
terms of his will. He lies here.[41]
The editors
of the RIB speculated that the last name Sita
may have derived from Sitas, a king
of “
Durnovaria (
It may be
seen from Tables 1 and 2, and from Figures 3-6, that E3b1a-M78 is
present in
Nevertheless,
the Romans were not absent from
Dark
(2002, pp. 202-207) has identified a Romano-British style settlement associated
with the kingdom of Strathclyde, very near the location of the western Scottish
haplotypes, appearing in Fig. 4. Some
evidence of Roman-style settlement also exists in the regions surrounding
modern
Summary
The
coalescence age (TMRCA) of E-V13, as calculated by Cruciani, et al. (2007), has
placed the expansion of this haplogroup from the
The E-M78
cluster E3b1α (phylogenetically equivalent to E-V13) was found by Peričić
(2005) to be at its highest frequency worldwide in the geographic region
corresponding to the Roman province of Moesia
Superior, a region that encompasses Kosovo, southern Serbia, northern
Macedonia and extreme northwestern Bulgaria today. The Roman invasion of CE 43, and the
subsequent occupation and settlement of Britain for nearly four centuries by
the Roman military, brought thousands of soldiers from the Balkan peninsula to
Britain as part of auxiliary units and, at least in some cases, as regular
legionnaires. By no later than the
mid-third century, retired Thracian soldiers and their families were living in
Deva (
With
E-M78 treated as a close approximation (85%) for the presence of E-V13, the
geographic association between the locations of E3b1a-M78 haplotypes and the
"Saxon Shore Forts" in southeastern Britain (especially in Norfolk)
may be significant, suggesting a possible link between the vicus populations of these forts and the current distribution of
these haplotypes. Similarly, the dense
settlement and long survival of sub-Roman Essex may be linked to the present
distribution of E3b1-M78 in southeastern
The lower
density of E3b haplotypes north of Hadrian's Wall may be explained by the more
limited Roman presence in that region, encompassing a period of approximately
100 years, rather than the 400+ years of Romano-British and sub-Roman society
south of the Wall. There is some indication of substantial Roman- and
sub-Roman-style settlement in Strathclyde and near
There
appears to be less doubt, after three published genetic surveys (Weale, 2002;
Capelli, 2003; Sykes, 2006), that the "
Some
relevant questions that must await better and more complete genetic sampling of
E3b1a2 haplotypes in Britain and western Europe include: (a) whether E-M78
(putatively E-V13) haplotypes from the Northern Wales/Cheshire geographic
cluster and from the southeastern England cluster are in fact from the same
population, originating in the Balkan peninsula, or whether their arrival times
and migration routes are substantially different; (b) what role (if any) J2-M12
has had in the Roman occupation and settlement of Britain; and, (c) could any
E3b haplotypes located in the Rhine river region also have been the result of
settlement and military occupation of Germania
Inferior by soldiers of Balkan origin?
It is
hoped that a large-scale genetic survey of all of
Acknowledgement
I wish to
thank Mr. Larry Jones, Esq., for his very helpful insights and commentary while
this manuscript was in preparation. Any
content, opinions or conclusions expressed, when not attributed directly to the
relevant source, are my own.
Electronic-Database Information
https://home.comcast.net/~hapest5/index.html
Haplogroup
Predictor Program
http://www.familytreedna.com/public/E3b/
Haplogroup
E3b Project
MapViewer
7, Golden Software
http://www.bloodoftheisles.net/results.html
http://www.roman-britain.org/main.htm
On-line
Roman Britain Resource
http://www.isogg.org/tree/index.html
ISOGG Y
Haplogroup Tree
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[1] "E3b" in
this and similar contexts refers to the E3b1-M35 clade and its component
subclades, collectively, unless otherwise specified.
[2] Oppenheimer (2006),
pp. 206-207.
[3] Ibid.
[4] Haplogroups
corresponding to E3b-M35 and J2-M172 in the Capelli study.
Address for
correspondence: stevenbird1000@hotmail.com
Received:
[5] Oxford Genetic Atlas Project
(OGAP) data is published online at: http://www.bloodoftheisles.net/results.html
[6] Irish data for Blood of the Isles was taken from
studies performed by the
[7]A current diagram of
the ten E3b1a-M78 subclades may be found at:
http://www.isogg.org/tree/
[8] Semino et al. (2004)
treated J-M12 and J-M102 as phylogenetically equivalent, as did
Peričić et al. (2005). Later
work has shown that J-M102 is a subgroup of J-M12 (ISOGG, 2007).
[9] The full data set from
both studies is available online at: http://www.gen.tcd.ie/molpopgen/resources.php
[10] These measures u
[11] An excellent map and discussion of
Sykes' OGAP regions may be found in K
Campbell (JoGG, Spring 2007, Fig. 1).
[12] The E3b1-M35 haplogroup is
identified by Sykes as “Eshu.”
[13] http://www.familytreedna.com/public/E3b/
[14] http://www.fluxus-engineering.com
[15] For all but a few haplotypes, the
Fluxus MJ network simply clarified and confirmed the haplogroup estimate
derived from the histograms and from a comparison with proved (SNP-tested) E3b
haplotypes. OGAP numbers 4022, A3059 and
A3165 were confirmed as E-M78 by their clustering within the MJ network. Haplotype 2745, which was estimated at first
(using the histograms) as a possible E-M34, was revised to M-78 based upon its
obvious clustering with three other M-78 haplotypes (A2243,
A2950, A1201); it was the only initial estimate that required revision.
[16] http://www.goldensoftware.com
[17] http://www.roman-britain.org/maps/british_tribes.htm.
18 Under this scenario,
members of other haplogroups besides E3b1-M78 would also have been displaced,
including, for example, R1b1-P25 and its subclades, but these haplotypes might
not be as easily distinguishable as "Romano-British," within the populations surveyed, as would be
those that are E3b1-M78.
19 Although the same
hypothesis may in fact apply to J2b-M12, the remainder of this study will focus
exclusively on E3b1-M35 and its component subclades, in particular E3b1a-M78
and E3b1a2-V13. More study of J2b-M12 and
its phylogeography is required before any firm conclusions may be made
concerning this clade. Such study,
although perhaps relevant, is beyond the scope of the present article.
[20] On the basis of the
positive identification of the α cluster of E-M78 with E-V13 by Cruciani,
it may be seen that E3b1a2, E-M78α
and E-V13 are phylogenetically equivalent to each other.
[21] This omission is
puzzling considering that Cruciani et al. (2007) acknowledged the findings of
Perečić et al. (2005) directly.
[22] See "
[23] The military action
that led to the conquest of
[24] Georgiev apparently
treated “Mysian” as a synonym for Moesian; this ethnonym should not be confused
with the Mysian people of
[25] RIB 2415.39.
[26] Item no.
1813.12-11.1-2, Department of Prehistory and Early
[27] RIB, v. II, 2401.5, p.
12.
[28] RIB, v. II, 2401.3, p. 8.
[29] The Bessi were a Thracian tribe who
lived in the
[30] For the purposes of this discussion,
"Thracian," "Dacian" and
"Bessus" (plural - "Bessi") are treated as
equivalent ethnonyms. The term
"Thracian" could have been applied to any of these groups, since the
descriptions "Bessus," "Dacian," and "Thracian"
were used by the Romans themselves rather indiscriminately.
[31] It is not clear from Mócsy's
description if he may be, in fact, referring to the same tombstone as
Carrington.
[32] Hogg was Secretary, Royal Commission
on Ancient and
[33] An interim excavation report is
available online at:
http://www.ancient-arts.org/Pentrwyn%20Bronze%20Age
%20Metalworking%20Site.htm#sitelo
[34] Sykes' region of "
[35] RIB 1909.
[36] RIB 1929b.
[37] Although the cavalry unit is
identified as Thracian, Serdica actually was located in the Roman
[38] It is often
speculated that the marker was thrown over during the Bouddican sack of
[39] RIB, no. 201. p. 66.
[41] RIB,
121.
[42] An excavation report
from Wessex Archaeology is available at:
http://www.wessexarch.co.uk/projects/dorset/dorchester_hospital
[43] The OGAP identified a small cluster
of three E3b samples found in the Shetland Islands, but since the haplotypes in
this one location were identical for the markers tested, they may well have
been all descended from a single ancestor, a "founder's effect."